RANGE EXTENSION OF TWO BRACHYURAN CRABS COLLECTED FROM THE AMAZON CONTINENTAL SHELF
DOI:
https://doi.org/10.32435/envsmoke-2026-0001Keywords:
Amazon Reefs, Bycatch, State of Amapá, Geographic distribution, New recordAbstract
The Amazon Continental Shelf (ACS) is composed of numerous benthic habitats, which shelter a high diversity of invertebrates, especially crustaceans. However, due to the complexity of ecosystems, knowledge of crustacean fauna is still incomplete. We herein report the northernmost Brazilian record of two brachyuran crab species, Teleophrys pococki Rathbun, 1924 (Mithracidae) and Euchirograpsus americanus A. Milne-Edwards, 1880 (Plagusiidae), associated with the Great Amazon Reef System (GARS) (State of Amapá), additionally, providing a description of their first gonopods (G1). Both male specimens E. americanus and T. pococki, were accidentally collected during commercial fishing operation of the red snapper Lutjanus purpureus (Poey, 1866) in the ACS (northern Brazil), at depths up to 70 m, in August 2023 and September 2025. The presence of both species in the GARS increases crustacean biodiversity in this ecosystem and highlights the need for further studies in the region to assess its true diversity and to develop management plans focused on species conservation.
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ALVES, D.F.; BARROS-ALVES, S.P.; TEIXEIRA, G.M.; COBO, V.J. Mithracinae (Decapoda: Brachyura) from the Brazilian coast: Review of the geographical distribution and comments on the biogeography of the group. Nauplius, v. 20, n. 1, p. 51–62, 2012. Available from: https://www.scielo.br/j/nau/a/ZxwV8fKNxv65FMVkKRxNH7J/?format=pdf&lang=en. Accessed on: 15 Sep 2025.
ALVES-JÚNIOR, F.A.; MARTINS, D.E.G.; KLAUTAU, A.G.C.M.; BARROS, A.P.C.; CINTRA, I.H.A. Pelagic Sargassum spp. blooms as an agent of marine litter transport along the Amazon continental shelf. Acta Fisheries and Aquatic Resources, v. 12, n. 2, p. 121–126, 2024a. Available from: https://doi.org/10.46732/actafish.2024.12.2.121-126
ALVES-JÚNIOR, F.A.; MARTINS, D.E.G.; CINTRA, I.H.A. Becoming a mother too early: The first record of the smallest ovigerous female of the deep-sea squat lobster Iridonida iris (A. Milne-Edwards, 1880) (Decapoda: Munididae). Pan-American Journal of Aquatic Science, v. 19, n. 2, p. 92–95, 2024b. Available from: https://doi.org/10.54451/PanamJAS.19.2.92
ALVES-JÚNIOR, F.A.; SANTANA, J.L.; ARAÚJO, M.S.L.C.; SOUZA-FILHO, J.F. First record of Euchirograpsus americanus A. Milne-Edwards, 1880 (Crustacea: Decapoda: Plagusiidae) from northeastern Brazil. Nauplius, v. 24, p. e2016023, 2016. Available from: https://doi.org/10.1590/2358-2936e2016023
BESCHIN, C.; BUSULINI, A.; TESSIER, G.; ZORZIN, R. I crostacei associati a coralli nell'Eocene inferiore dell'area di Bolca (Verona e Vicenza, Italia nordorientale). Memorie del Museo Civico di Storia Naturale di Verona, serie 2, Sezione Scienze della Terra 9, 189 p., 2016. Available from: https://www.valdalponeheritage.it/ATS_Documenti/2016_I%20Crostacei%20associati%20a%20coralli.pdf. Accessed on: 20 Oct 2025.
CANO, G. Viaggio della R. Corvetta Vettor Pisani attorno al globo. – Crostacei Brachiuri ed Anomuri. Bollettino della Società di naturalisti in Napoli, v. 3, p. 169–268, 1889. Available from: https://www.marinespecies.org/aphia.php?p=sourcedetails&id=391827. Accessed on: 15 Oct. 2025.
CARMONA-SUÁREZ, C.; POUPIN, J. Majoidea crabs from Guadeloupe Island, with a documented list of species for the Lesser Antilles (Crustacea, Decapoda, Brachyura, Majoidea). Zoosystema, v. 38, n. 3, p. 353–387, 2016. Available from: https://doi.org/10.5252/z2016n3a5
CASTAÑO, N.C.; CAMPOS, N.H. Los cangrejos Araña (Decapoda: Brachyura: Majoidea) del Caribe colombiano. Biota Colombiana, v. 4, n. 2, p. 261–269, 2003. Available from: https://www.redalyc.org/articulo.oa?id=49140208 . Accessed on: 17 Sep 2025.
COELHO, P.A.; ALMEIDA, A.O.; BEZERRA, L.E.A. Checklist of the marine and estuarine Brachyura (Crustacea: Decapoda) of the northern and northeastern Brazil. Zootaxa, v. 1956, n.1, p. 1–58, 2008. Available from: https://doi.org/10.11646/zootaxa.1956.1.1
CORBARI, L.; MARTIN-LEFÈVRE, P.; SABROUX, R.; SOUBZMAIGNE, S. Crustacés. In: PASCAL, O.; TOUROULT, J.; BOUCHET, P. (Eds.). Expédition La Planète Revisitée Guyane 2014–15: Synthèse des Premiers Résultats. Muséum national d’Histoire naturelle. p. 162–174, 2015. Available from https://inpn.mnhn.fr/docs-web/docs/download/249749. Accessed on: 17 Sep 2025.
DANA, J.D. Crustacea Grapsoidea, (Cyclometopa, Edwardsii): Conspectus Crustacearum quæ in Orbis Terrarum circumnavigatione, Carolo Wilkes e classe Reipublicæ Fœderatæ Duce, lexit et descriptsit J. D. Dana. Proceedings of the Academy of Natural Sciences of Philadelphia, v. 5, p. 247–254. Available from: https://www.biodiversitylibrary.org/part/22775. Accessed on: 05 Nov 2025.
DecaNet (World List of Decapoda). DecaNet. Mithracidae MacLeay, 1838. 2025a. Available from: https://www.decanet.info/aphia.php?p=taxdetails&id=439056. Accessed on: 12 Nov 2025.
DecaNet (World List of Decapoda). DecaNet. Plagusiidae Dana, 1851. 2025b. Available from: https://www.decanet.info/aphia.php?p=taxdetails&id=439056. Accessed on: 12 Nov 2025.
DRAKE, T.W.; HEMINGWAY, J.D.; KUREK, M.R.; PEUCKER-EHRENBRINK, B.; BROWN, K.A.; HOLMES, R.M.; VALIER, G.; MOURA, J.M.S; MITSUYA, M.; WASSENAAR, L.I.; SIX, J.; SPENCER, R.G.M. The pulse of the Amazon: Fluxes of dissolved organic carbon, nutrients, and ions from the world's largest river. Global Biogeochemical Cycles, v. 35, n. 4, p. e2020GB006895, 2021. Available from: https://doi.org/10.1029/2020GB006895
FLEEGER, S.T.D.J.; COWAN-JR., J.H.; PASCAL, P-.Y. What Is the Relative Importance of Phytoplankton and Attached Macroalgae and Epiphytes to Food Webs on Offshore Oil Platforms? Marine and Coastal Fisheries: Dynamics, Management, and Ecosystem Science, v. 5, p. 53–64, 2013. Available from: https://doi.org/10.1080/19425120.2013.774301
FUJITA, Y.; NARUSE, T. A new genus and species of a submarine cave crab of the family Plagusiidae Dana, 1851 (Crustacea: Brachyura: Grapsoidea) from Okinawa Island, Ryukyu Islands, southwestern Japan. Zootaxa, v. 5410, n. 3, p. 408–418, 2024. Available from: https://doi.org/10.11646/zootaxa.5410.3.8
GARTH, J.S. Marine Biological Investigations in the Bahamas 19. Decapoda Brachyura. Sarsia, v. 63, n. 4, p. 317–333, 1978. Available from: https://doi.org/10.1080/00364827.1978.10411352
GOUVÊA, E.P. A carcinofauna do litoral rochoso de Salvador, BA, e alguns aspectos ecológicos. Ciência e Cultura, v. 38, n. 2, p. 346–354, 1986. Available from: http://memoria.bn.gov.br/docreader/003069/45802. Accessed on: 08 Oct 2025.
GUINOT, D. A new genus of the family Plagusiidae Dana, 1851, close to Plagusia Latreille, 1804 (Crustacea, Decapoda, Brachyura). Zootaxa, v. 1498, n. 1, p. 27–33, 2007. Available from: https://doi.org/10.11646/zootaxa.1498.1.2
HENDRICKX, M.E. On a small collection of decapod crustaceans (Malacostraca, Decapoda) from the Galapagos Islands, collected by P. H. Vercammen-Grand-Jean, in 1964. Crustaceana, v. 83, n. 8, p. 1009–1015, 2010. Available from: https://doi.org/10.1163/001121610X502902
HOLTHUIS, L.B. A small collection of Decapod Crustacea from Galapagos Islands. Spedizione L. Mares-G.R. T. S., Florence, Italy, 1979. Available from: https://scispace.com/pdf/a-small-collection-of-decapod-crustacea-598sdro3bn.pdf. Accessed on: 13 Nov 2025.
KLAUTAU, A.G.C.M.; CORDEIRO, A.P.B.; CHAGAS, R.A.; SANTOS, W.C.R.; MARCENIUK, A.P.; NÓBREGA, P.S.V.; MARTINELLI-LEMOS, J.M.; CINTRA, I.H.A.; CASTRO, N.S.S.; BASTOS, C.E.M.C.; ALVES-JÚNIOR, F.A.; PINHEIRO, L.C.; BENTES, B. Biodiversity hotspots and threatened species under human influence in the Amazon continental shelf. Scientific Reports, v. 15, p. 26681, 2025. Available from: https://doi.org/10.1038/s41598-025-11261-x
KÜTZING, F.T. (1842 '1841'). Ueber Ceramium Ag. Linnaea, v. 15, n. 6, p. 727–746, 1841. Available from: https://www.algaebase.org/search/bibliography/detail/?biblio_id=11288. Accessed on: 15 Nov 2025.
KÜTZING, F.T. Phycologia Germanica, d.i. Deutschlands Algen in bündigen Beschreibungen. Nebst einer Anleitung zum Untersuchen und Bestimmen dieser Gewächse für Anfänger. Nordhausen, 1845, 340 pp., Available from: https://doi.org/10.5962/bhl.title.13687
LAMARCK, J.B.M. Histoire naturelle des animaux sans vertèbres, Tome troisième, Paris, Deterville/Verdière, 586 p., 1816. Available from: https://doi.org/10.5962/bhl.title.12712
LATREILLE, P.A. Tableau méthodique des crustacés. Classe septième. Crustacés (1), Crustacea. Nouveau Dictionnaire d'Histoire Naturelle, Appliquée aux Arts, Principalement à l'Agriculture et à l'Economie Rurale et Domestique: par une Société de Naturalistes et d'Agriculteurs: avec des figures tirées des trois Règnes de la Nature. v. 24, p. 123–127, 1804. Available from: https://doi.org/10.5962/bhl.title.151939
LÓPEZ, M.M.; SOLANO, O.D. Estado poblacional de Echinometra lucunter (Echinoida: Echinometridae) y su fauna acompañante en el litoral rocoso del Caribe Colombiano. Revista de Biologia Tropical, v. 53, n. 3, p. 291–297, 2005. Available from: https://tropicalstudies.org/rbt/attachments/suppls/sup53-3%20echinoderm/19-MONROY-Asp.pdf. Accessed on: 05 Nov 2025.
MACLEAY, W.S. Illustrations of the Annulosa of South Africa. On the Brachyurous Decapod Crustacea. Brought from the Cape by Dr. Smith. In: Smith, A. (Ed.), Illustrations of the zoology of South Africa: consisting chiefly of figures and descriptions of the objects of natural history collected during an expedition into the interior of South Africa, in the years 1834, 1835, and 1836, fitted out by “The Cape of Good Hope Association for Exploring Central Africa”: together with a summary of African zoology, and an inquiry into the geographical ranges of species in that quarter of the globe. London: Authority of the Lords Commissioners of Her Majesty's Treasury, 1838. Vol. 5. p. 53–71. Available from: https://www.biodiversitylibrary.org/item/41545. Accessed on: 05 Nov 2025.
MANNING, R.B.; HOLTHUIS, L.B. West African brachyuran crabs (Crustacea: Decapoda). Smithsonian Contributions to Zoology, v. 306, p. 1–379, 1981. Available from: https://doi.org/10.5479/si.00810282.306
MARTINS, D.E.G.; ALVES-JÚNIOR, F.A.; CINTRA, I.H.A. New record of the swimming crab Portunus sayi (Gibbes, 1850) (Decapoda: Portunidae) from the Amazon continental shelf. Environmental Smoke, v. 8, p. es20250003, 2025. Available from: https://doi.org/10.32435/envsmoke-2025-0003
MELO, G.A.S. Manual de identificação dos Brachyura (caranguejos e siris) do litoral brasileiro. Editora Plêiade, São Paulo, 603 pp, 1996. Available from: https://repositorio.usp.br/item/000921427. Accessed on: 10 Oct. 2025.
MELO, G.A.S. Malacostraca-Eucarida. Brachyura. Oxyrhyncha and Brachyrhyncha. p. 455–515. In: YOUNG, P.S. (ed). Catalogue of Crustacea of Brazil. Rio de Janeiro, Museu Nacional, Série Livros 6, 1998. Available from: https://www.scielo.sa.cr/scieloOrg/php/reflinks.php?refpid=S0034-7744200800030002100053&lng=es&pid=S0034-77442008000300021. Accessed on: 18 Oct. 2025.
MILNE-EDWARDS, A. Reports on the results of dredging, under the supervision of Alexander Agassiz, in the Gulf of Mexico and in the Caribbean Sea, 1877, 1878, 1879, by the U.S. Coast Survey Steamer “Blake”, Lieut-Commander C.D. Sigsbee, U.S.N., and Commander J.R. Bartlett, U.S.N., Commanding. VIII. Études préliminaires sur les Crustacés, 1re Partie. Bulletin of the Museum of Comparative Zoology at Harvard College, v. 8, n. 1, p. 1–68, 1880. Available from: https://biostor.org/reference/106168. Accessed on: 05 Nov 2025.
MILNE-EDWARDS, A.; BOUVIER, E.L. Crustacés décapodes provenant des campagnes du yacht l’Hirondelle (1886, 1887 et 1888). Première partie: Brachyures et Anomoures. Résultats des Campagnes Scientifique Monaco, v. 7, p. 1–112, 1894. Available from: https://www.biodiversitylibrary.org/item/31107. Accessed on: 15 Oct 2025.
MILNE-EDWARDS, H. Mémoire sur la famille des ocypodiens. Annales des Sciences Naturelles, v. 20, p. 163–228, 1853. Available from: https://biostor.org/reference/160231. Accessed on: 16 Oct 2025.
MOURA, R.L.; AMADO-FILHO, G.M.; MORAES, F.C.; BRASILEIRO, P.S.; SALOMON, P.S.; MARQUES, M.M.; BASTOS, A.C.; ALMEIDA, M.G.; SILVA-JÚNIOR, J.M.; ARAÚJO, B.F.; BRITO, F.P.; RANGEL, T.P.; OLIVEIRA, B.C.V.; BAHIA, R.G.; PARANHOS, R.P.; DIAS, R.J.S.; SIEGLE, E.; FIGUEIREDO-JÚNIOR, A.G.; PEREIRA, R.C.; LEAL, C.V.; HAJDU, E.; ASP, N.E.; GREGORACCI, G.B.; NEUMANN-LEITÃO, S.; YAGER, P.L.; FRANCINI-FILHO, R.B.; FROÉS, A.; CAMPEÃO, M.; SILVA, B.S.; MOREIRA, A.P.B.; OLIVEIRA, L.; SOARES, A.C.; ARAÚJO, L.; OLIVEIRA, N.L.; TEIXEIRA, J.B.; VALLE, R.A.B.; THOMPSON, C.C.; REZENDE, C.E.; THOMPSON, F.L. An extensive reef system at the Amazon River mouth. Science Advances, v. 2, n. 4, p. 1–11, 2016. Available from: https://doi.org/10.1126/sciadv.1501252
NG, P.K.L.; GUINOT, D.; DAVIE, P.J.F. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology, v. 17, n. 1, p. 1–286, 2008. Available from: https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/11/app/uploads/2017/04/s17rbz.pdf. Accessed on: 03 Nov 2025.
NOGUEIRA, M.M.; NEVES, E.; JOHNSSON, R. Effects of habitat structure on the epifaunal community in Mussismilia corals: does coral morphology influence the richness and abundance of associated crustacean fauna?. Helgoland Marine Research, v. 69, p. 221–229, 2015. Available from: https://doi.org/10.1007/s10152-015-0431-x
PORTELL, R.W.; COLLINS, J.S.H. Decapod crustaceans of the Lower Miocene Montpellier Formation, White Limestone Group of Jamaica. Cainozoic Research. v. 3, n. 1–2, p. 109–126, 2004. Available from: https://natuurtijdschriften.nl/pub/541705/CR2004003001007.pdf. Accessed on: 15 Oct 2025.
POUPIN, J. Faune marine profonde des Antilles françaises: récoltes du navire Polka faites en 1993. Paris: ORSTOM, 1994. Collection Études et Thèses, 1994. Available from: https://horizon.documentation.ird.fr/exl-doc/pleins_textes/pleins_textes_2/etudes_theses/39603.pdf. Accessed on: 16 Sep 2025.
POUPIN, J. Les Crustacés décapodes des Petites Antilles. Avec de nouvelles Observations pour Saint-Martin, la Guadeloupe et la Martinique. Muséum National d’Histoire Naturelle, Patrimoines naturels 77, 2018. Available from: https://sciencepress.mnhn.fr/sites/default/files/documents/en/cpn77-listedoc.pdf. Accessed on:12 Nov 2025.
POUPIN, J. A documented checklist of the Crustacea (Stomatopoda, Decapoda) of the southern Guianas (Guyana, Suriname, French Guiana, Brazil Amapá). European Journal of Taxonomy, v. 954, p. 1–197, 2024. Available from: https://doi.org/10.5852/ejt.2024.954.2653
RATHBUN, M.J. The decapod and stomatopod Crustacea. I. Results of the Branner-Agassiz Expedition to Brazil. Proceedings of the Washington Academy of Sciences, v. 2, p. 133–156, 1900. Available from: https://www.biodiversitylibrary.org/page/8872151#page/172/mode/1up. Accessed on: 06 Nov 2025.
RATHBUN, M.J. Investigations of the Aquatic Resources and Fisheries of Porto Rico by the United States Fish Commission Steamer Fish Hawk in 1899. The Brachyura and Macrura of Porto Rico. Bulletin of the United States Fish Commission, v. 20, p. 1–127, 1901. Available from: https://www.decanet.info/aphia.php?p=sourcedetails&id=125904. Accessed on: 12 Nov 2025.
RATHBUN, M.J. The grapsoid crabs of America. United States National Museum Bulletin, v. 97, p. 1–461, 1918. Available from: https://repository.si.edu/items/239cd4de-44e5-424d-93b9-3d72ed7813dd. Accessed on: 8 Nov 2025.
RATHBUN, M.J. New species and subspecies of spider crabs. Proceedings of the United States National Museum. v. 64, n. 2504, p. 1–5, 1924. Available from: https://repository.si.edu/server/api/core/bitstreams/5a41306b-fbab-40f8-bb32-2db44a44f9f7/content. Accessed on: 03 Nov 2025.
RATHBUN, M.J. The spider crabs of America. United States National Museum Bulletin, v. 129, p. 1–613, 1925. Available from: https://repository.si.edu/items/d54c874d-86aa-4a49-8481-188e9442518b. Accessed on: 06 Nov 2025.
ROTH, A.W. Catalecta Botanica (Quibus plantae novae et minus cognitae describuntur atque illustrantur). Fasc. 1. Lipseae., 244 p., 1797. Available from: https://www.biodiversitylibrary.org/page/56933819. Accessed on: 11 Nov 2025.
SCHUBART, C.D.; CUESTA, J.A. Phylogenetic relationships of the Plagusiidae Dana, 1851 (Brachyura), with description of a new genus and recognition of Percnidae Stevcic, 2005, as an independent family. In: CASTRO, P.; DAVIE, P.J.F.; NG, P.K.L.; FORGES, B.R. Studies on Brachyura: a Homage to Danièle Guinot. Crustaceana Monographs 11. Leiden: Brill. p. 279–300, 2010. Available from: https://decapoda.nhm.org/pdfs/31482/31482.pdf. Accessed on: 12 Oct 2025.
STIMPSON, W. Sketch of a revision of the genera of Mithracidae. The American Journal of Science and Arts, Series 2, v. 29, n. 85, p. 132–133, 1860. Available from: https://archive.org/details/sketchofrevision00stim/page/132/mode/2up. Accessed on: 28 Oct 2025.
STIMPSON, W. Preliminary report on the Crustacea dredged in the Gulf Stream in the Straits of Florida, by L. F. de Pourtales, Assist. U. S. Coast Survey. Part I. Brachyura. Bulletin of the Museum of Comparative Zoölogy, v. 2, n. 2, p. 109–160, 1871. Available from: https://www.biodiversitylibrary.org/page/6313618#page/125/mode/1up. Accessed on: 18 Oct 2025.
TAVARES, M.; MENDONÇA Jr., J.B. Brachyuran crabs (Crustacea, Decapoda) from the remote oceanic Archipelago Trindade and Martin Vaz, South Atlantic Ocean. Zootaxa, v. 5146, n. 1, p. 1–129, 2022. Available from: https://doi.org/10.11646/zootaxa.5146.1.1
TÜRKAY, M. Zur Kenntnis der Gattung Euchirograpsus mit Bemerkungen zu Brachygrapsus und Litocheira (Crustacea, Decapoda). Senckenbergiana biologica, v. 56, n. 1, p. 103–132, 1975. Available from: https://decapoda.nhm.org/pdfs/12714/12714.pdf. Accessed on: 14 Oct 2025.
TÜRKAY, M. Zwei neue Grapsiden-Arten aus dem Indopazifik (Euchirograpsus madagascariensis n. sp. und Miersiograpsus australiensis n. sp.) mit Einführung von Miersiograpsus n. gen. (Crustacea: Decapoda: Grapsidae). Senckenbergiana biologica, v. 59, n. 1–2, p. 133–141, 1978. Available from: https://decapoda.nhm.org/pdfs/23688/23688.pdf. Accessed on: 11 Oct 2025.
VALE, N.F.; BRAGA, J.C.; MOURA, R.L.; SALGADO, L.T.; MORAES, F.C.; KAREZ, C.S.; CARVALHO, R.T.; SALOMON, P.S.; MENANDRO, P.S.; AMADO-FILHO, G.M.; BASTOS, A.C. Distribution, morphology and composition of mesophotic ‘reefs’ on the Amazon continental margin. Marine Geology, v. 447, p. 106779, 2022. Available from: https://doi.org/10.1016/j.margeo.2022.106779
VERA-CARIPE, J.; DIAZ, O.; LIRA, C.; BOLAÑOS, J. Crustáceos decápodos asociados a Tripneustes ventricosus (Lamarck, 1816) (Echinodermata; Echinoidea) de la isla La Borracha, parque nacional Mochima, Venezuela. Boletín del Instituto Oceanográfico de Venezuela, v. 56, n. 1, p. 61–68, 2017. Available from: http://saber.udo.edu.ve/index.php/boletiniov/article/view/2644/1566. Accessed on: 04 Nov 2025.
VERRILL, A. E. Variations and nomenclature of Bermudian, West Indian and Brazilian reef corals, with notes on various Indo-Pacific corals. Transactions of the Connecticut Academy of Arts and Sciences, v. 11, p. 63–168, 1901. Available from: https://www.biodiversitylibrary.org/page/13464046. Accessed on: 25 Oct 2025.
WEBER, S.C.; CARPENTER, E.J.; COLES, V.J.; YAGER, P.L.; GOES, J.; MONTOYA, J.P. Amazon River influence on nitrogen fixation and export production in the western tropical North Atlantic. Journal of Oceanology and Limnology, v. 62, p. 618–631, 2017. Available from: https://doi.org/10.1002/lno.10448
WILSON, B.M.; POHLE, G.W. Northern range expansion of the American talon crab, Euchirograpsus americanus A. Milne-Edwards, 1880 (Decapoda, Grapsoidea, Plagusiidae), to the Bay of Fundy, Canada. Crustaceana, v. 89, n. 2, p. 163–173, 2016. Available from: https://www.jstor.org/stable/43965981. Accessed on: 18 Oct 2025.
WINDSOR, A.M.; FELDER, D.L. Molecular phylogenetics and taxonomic reanalysis of the family Mithracidae MacLeay (Decapoda: Brachyura: Majoidea). Invertebrate Systematics, v. 28, p. 145–173, 2014. Available from: https://doi.org/10.1071/IS13011
ZARIQUIEY-ALVAREZ, R. Crustáceos decápodos ibéricos. Investigación Pesquera, Barcelona, v. 32, p. 1–510, 1968. Available from: https://decapoda.nhm.org/pdfs/10992/10992.pdf. Accessed on: 22 Oct 2025.
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